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The Ancestors of the Diptera

Abstract

IN a previous communication to NATURE some years ago (May 18, 1929) I gave a short description of a remarkable genus Permotipula, represented by a complete forewing from Warner's Bay, N.S.W., and of Upper Permian Age. Although very primitive by comparison with existing types of Diptera, this wing was regarded by me as being definitely Tipuloid and lying well within the order. I added the following remarks: ” The wing is of the greatest interest, because any student of venation would classify it as dipterous and nothing else, and yet we do not know whether the insect to which it belonged had four wings or only two! Also, it is the oldest known dipterous type of wing by many millions of years.”

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TILLYARD, R. The Ancestors of the Diptera. Nature 139, 66–67 (1937). https://doi.org/10.1038/139066a0

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