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Protein-mediated membrane fusion is an essential, common denominator for viral infection, fertilization, intracellular protein trafficking, neurotransmission, hormone release, developmental cell-cell fusion, and autophagy. This Collection will shed light on the mechanisms of membrane fusion and will include studies in areas such as: the structure and the function of fusogens and auxiliary regulators, lipid dynamics, bilayer remodelling, fusion intermediates, and fusion pore dynamics.