Amphibians hint that number skills evolved early.
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Uller, C., Jaeger, R., Guidry, G. & Martin, C. Salamanders (Plethodon cinereus) go for more: rudiments of number in an amphibian. Animal Cognition, published online, doi:10.1007/s10071-003-0167-x (2003).
Nieder, A., Freedman, D. J. & Miller, E. K. Representation of the quantity of visual items in the primate prefrontal cortex. Science, 297, 1708 - 1711, (2003).
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Hoag, H. Salamanders can do maths. Nature (2003). https://doi.org/10.1038/news030428-19