Water molecules and DNA conformation are now recognized as ingredients which can influence both the affinity and specificity of protein/DNA complexes.
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Arbuckle, N., Luisi, B. A recipe for specificity. Nat Struct Mol Biol 2, 341–346 (1995). https://doi.org/10.1038/nsb0595-341
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DOI: https://doi.org/10.1038/nsb0595-341
