Ancient DNA contains clues about complexion.
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References
Lalueza-Fox, C. et al. Science doi:10.1126/science.1147417 (2007).
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Ledford, H. Some Neanderthals were red-heads. Nature (2007). https://doi.org/10.1038/news.2007.197
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DOI: https://doi.org/10.1038/news.2007.197