A cell's shape changes as it moves along a surface. The forward-thinking cytoskeletal elements are all for progress, but the conservative cell membrane keeps them under control by physically opposing their movement.
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Haugh, J. Cells get in shape for a crawl. Nature 453, 461–462 (2008). https://doi.org/10.1038/453461a
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DOI: https://doi.org/10.1038/453461a