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Acknowledgments.-The sequential model fitting was carried out on the KDF9 computer of the University of Birmingham.
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Perkins, J., Jinks, J. Detection and estimation of genotype-environmental, linkage and epistatic components of variation for a metrical trait. Heredity 25, 157–177 (1970). https://doi.org/10.1038/hdy.1970.22
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DOI: https://doi.org/10.1038/hdy.1970.22
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