Table 3: Relatedness* among males, and males and infants, in the attacking groups.

From: Observations of severe and lethal coalitionary attacks in wild mountain gorillas

GroupNumber of dyads that were…Mean relatedness coefficient for…
UnrelatedFather- sonFull siblingsMaternal siblingsPaternal siblingsAll males 8+ yearsAll males 8+ years to infants
2004 (Group Beetsme)7610140.250.17
2010 (Group Pablo)101120450.25Unknownx
2013 (Group Titus)000070.25Unknownt
  1. *Only parent-offspring and sibling relationships are considered here. Relatedness amongst all animals in this small (n = ~48042), closed population is quite high. Paternity data from ref. 63.
  2. xPaternity undetermined at the time of publication for 5 of 6 infants. Male CAN, who sired 11 of the 12 natal males, also sired 3 of the infants’ mothers, who were therefore (minimally) grandoffspring or half nieces/nephews to 12 of the males. CAN’s maternal brother sired the infant whose paternity was known. A fifth infant had an adult maternal brother in the group.
  3. tPaternity undetermined at the time of publication for the group’s one infant. The infant’s mother was neither paternally nor maternally related to any of the males.