Scientific Reports 6: Article number: 22147; published online: 26 February 2016; updated: 14 November 2016

The original version of this Article contained errors in the affiliations.

‘Institute of Basic Medical Sciences, National Cheng Kung University, Tainan, Taiwan’

was incomplete, and now reads:

‘Institute of Basic Medical Sciences, College of Medicine, National Cheng Kung University, Tainan, Taiwan’

This error has now been corrected in the HTML and PDF versions of this Article.