Scientific Reports 5: Article number: 14985; published online: 08 October 2015; updated: 14 November 2016

The original version of this Article contained errors in the affiliations.

‘Department of Cell Biology and Anatomy, National Cheng Kung University, North District, Tainan City, Taiwan’

was incomplete, and now reads:

‘Department of Cell Biology and Anatomy, College of Medicine, National Cheng Kung University, Tainan, Taiwan, ROC’

This error has now been corrected in the HTML and PDF versions of this Article.