a-c, Output filters for each feedback cue (see Fig. 1b) that predict the emission of each song type for a given state. ‘No song’ filters are not shown as these are fixed to be constant, and song type filters are in relation to these values (see Methods). Heavy line represents mean, shading represents SEM. d-e, Sign of filter for each emission filter shows the same feature can be excitatory or inhibitory depending on the state.