Correction to: Nature Genetics, published online 25 February 2019.

In the version of this article initially published, the y-axis labels in Fig. 5 read ‘signed log10[FDR]’ but should have read ‘–log10[FDR]’, and the figure legend incorrectly indicated that directions of effect are shown in Fig. 5 rather than Supplementary Table 25. The errors have been corrected in the HTML and PDF versions of the article.