Correction to: Nature Genetics https://doi.org/10.1038/s41588-018-0339-x, published online 11 February 2019.

In the version of this article initially published, in the key of Fig. 4, the two labels reading ‘FDR < 0’ should have read ‘FDR < 0.1’ and ‘FDR > 0.1’. The errors have been corrected in the HTML and PDF versions of the article.