Correction to: Nature Biotechnology, published online 3 December 2019.

In the version of this article initially published, an affiliation for author Zheng Wang was missing. This affiliation should appear first; it reads “School of Basic Medicine, Qingdao University, Qingdao, China.” Also, as a result of a publisher error, the matrix variable P was replaced by tbfP in equation (4), the matrix variable Y was formatted italic rather than bold in the first two instances of Ylandmarks, and, in the PDF version, some instances of the variables P, x, y, p, B, J, Q, K, Y and v were formatted as non-italic plain text instead of non-italic bold. The errors have been corrected in the HTML and PDF versions of the article.