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  1. Department of Medicine, Johns Hopkins University School of Medicine, Baltimore, MD, USA

    • Lynn N. Bertagnolli
    • , Jennifer A. White
    • , Francesco R. Simonetti
    • , Subul A. Beg
    • , Jun Lai
    • , Alexandra J. Murray
    • , Annukka A. R. Antar
    • , Robert F. Siliciano
    • , Gregory M. Laird
    •  & Janet D. Siliciano
  2. Howard Hughes Medical Institute, Baltimore, MD, USA

    • Subul A. Beg
    • , Jun Lai
    •  & Robert F. Siliciano
  3. The Wistar Institute, Philadelphia, PA, USA

    • Costin Tomescu
    •  & Luis J. Montaner
  4. Department of Molecular Microbiology and Immunology, Johns Hopkins Bloomberg School of Public Health, Baltimore, MD, USA

    • Hao Zhang
    •  & Joseph B. Margolick
  5. Division of HIV, Infectious Diseases and Global Medicine, University of California, San Francisco, CA, USA

    • Rebecca Hoh
    •  & Stephen G. Deeks
  6. Division of Infectious Diseases, Department of Medicine, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA, USA

    • Pablo Tebas


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L.N.B., J.A.W., G.M.L., F.R.S., R.F.S. and J.D.S. designed experiments. S.A.B., C.T. and L.J.M. obtained samples. L.N.B., J.A.W., S.A.B., G.M.L., F.R.S., J.L., A.J.M., A.A.R.A. and J.D.S. performed experiments. F.R.S., H.J. and J.B.M. performed cell sorting. L.N.B., J.A.W., F.R.S., A.J.M., A.A.R.A., R.F.S. and J.D.S. analysed the data and wrote the manuscript.

Competing interests

Declared none.

Corresponding author

Correspondence to Robert F. Siliciano.

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  1. Supplementary Information

    This file contains Supplementary Tables 1-3, Supplementary Figures 1-2 and the Supplementary Methods.

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