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Acknowledgements

I am grateful to A. Simpson, M. Leger, M. O’Malley, J. McCutcheon, T. Richards, F. Doolittle, L. Eme and A. Inkpen for discussion and critical review of the text. Financial support for this work comes from the Canada Research Chairs programme, the Natural Sciences and Engineering Research Council and the Canadian Institutes of Health Research.

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Correspondence to Andrew J. Roger.

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