*The assignment of apoproteins (LhcbM1-3) in M-LHCII or L-LHCII is tentative due to insufficient resolution of the local map features. The assignment of Chl a/b and carotenoids in M/L-LHCIIs is deduced from those in S-LHCII of the C2S2 complex. BCR, β-carotene; Lut, lutein; Neo, neoxanthin; Vio, violaxanthin; MGDG, monogalactosyl diacylglycerol; SQDG, sulfoquinovosyl diacylglycerol; PG, phosphatidylglycerol; DGDG, digalactosyl diacylglycerol; BCT, bicarbonate ion; LMU, dodecyl-α- D-maltoside; PL9, plastoquinone 9. When the contents are different in two structures, the numbers are separated by ‘/' (left, stacked C2S2; right, C2S2M2L2). The dash symbols indicate that the subunits or cofactors are not observed in the structure.