Correction to: Nature Communications https://doi.org/10.1038/ncomms10523, published online 27 January 2016.

The original version of this Article contained an error in author affiliations. Yejie Shi was incorrectly associated with China-America Institute of Neuroscience, Xuanwu Hospital, Capital Medical University, Beijing 100053, China, instead of Geriatric Research, Educational and Clinical Center, Veterans Affairs Pittsburgh Health Care System, Pittsburgh, PA 15261, USA. This has not been corrected in both the PDF and HTML versions of the Article.