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Publisher’s note: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.
The authors would like to thank Barry Zee, Tobin Gramyk, George Murphy and Christine Lian for critical reading of the manuscript and insightful comments, Guoming Shi, Jia Fan, Fei Lan and Lingchun Kong for their help and advice in experimental design, Anna Payne-Tobin Jost and Jennifer Waters of NIC at Harvard for the supports in imaging, and Santa Cruz Biotechnology, Inc. for providing antibodies. This work was supported by NIH grants GM112062 and CA194302 to Y.G.S.