Scottish millipede represents earliest known air-breathing animal.
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Wilson, H. M. & Anderson, L. I. Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland. Journal of Paleontology, 78, 169 - 184, (2004).
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Hopkin, M. Fossil find breaks age record. Nature (2004). https://doi.org/10.1038/news040126-1