Ancient male fish may have had a long, articulated organ for mating.
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References
Long, J., Trinajstic, K. & Johanson, Z. Nature 457, 1124–1127 (2009).
Long, J. A., Trinajstic, K., Young, G. C. & Senden, T. Nature 453, 650–652 (2008).
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Cressey, D. Fossils reveal early evidence for penetrative sex. Nature (2009). https://doi.org/10.1038/news.2009.122
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DOI: https://doi.org/10.1038/news.2009.122