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This 300-Myr example actually demonstrates the utility of our method. To see this, consider mantle flow beneath a supercontinent covering one-third of the globe: mantle upwelling is expected beneath the opposing ocean’s spreading ridges, but mantle downwelling occurs neither opposite to this upwelling (as for dipole flow) nor in bands 90° away (as for quadrupole flow), but instead associates with subduction occurring between these two locations on the supercontinent’s periphery. Return flow from this downwelling should drive upwellings beneath both the supercontinent and the oceanic plates. Indeed, upwelling is expected beneath a supercontinent that will soon disperse3,4.

Thus, we expect strong upwelling beneath the oceanic plates and weaker upwelling beneath the supercontinent; such a flow field is described by a combination of dipole and quadrupole flow fields that partially cancel beneath the supercontinent. This pattern is exactly predicted by the net characteristics (or spherical harmonics) of surface plate motions: the 300-Myr analysis of ref. 1 shows weak divergence within the supercontinent, indicating underlying upwelling. Thus, the lifetime of the two antipodal upwellings in the mantle may extend beyond the 250 Myr that we demonstrated in our original Letter2. More importantly, this analysis1 demonstrates the importance of using only the longest-wavelength components of plate motions to visualize the underlying mantle flow patterns. By including shorter-wavelength spherical harmonic degrees, Rudolph and Zhong have incorporated the influence of regional and local tectonics into their interpretation1; doing this obscures the underlying mantle flow patterns that are only apparent at the longest wavelengths2.

We agree with the Comment1 that quadrupole stability alone does not prove long-term stability of the LLSVP regions, and that additional constraints from “robust observations” are necessary. Indeed, the locations of large igneous provinces and kimberlites have been shown to source from the margins of two antipodal LLSVPs5, and would arise above a cold downwelling on the African side if the degree-1 interpretation of ref. 1 is correct. Furthermore, the 300-Myr plate motion example1 is based on a study6 that does not control for palaeolongitude or true polar wander, so it is unclear how surface features are related to LLSVP locations. Their portrayal of Pangaea as a stable coherent polygon additionally ignores much of the tectonic complexity of that supercontinent’s evolution7. These problems illustrate the importance of using a carefully reconstructed model of past plate motions when attempting to use “net characteristics” to constrain LLSVP stability.