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Nature volume 458, page E5 (19 March 2009) | Download Citation



Replying to: J. H. Geisler & J. M. Theodor Nature 458, 10.1038/nature07776 (2009)

The analysis of Geisler and Theodor1 confirms our main phylogenetic result2, that raoellids are, or include, the sister group to cetaceans. Their study expands on our findings by inferring that hippopotamids are the sister group to the combined raoellid–cetacean clade, whereas our paper had explicitly stated that our data could not address the position of the extant artiodactyl families.

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    & Hippopotamus and whale phylogeny. Nature 458 10.1038/nature07776 (2009)

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    , , & in The Evolution of Artiodactyls (eds Prothero, D. R. & Foss, S. E.) 19–31 (Johns Hopkins Univ. Press, 2007)

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    & Impact of increased character sampling on the phylogeny of Cetartiodactyla (Mammalia): combined analysis including fossils. Cladistics 24, 397–442 (2008)

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  1. *Department of Anatomy, Northeastern Ohio Universities College of Medicine, Rootstown, Ohio 44272, USA.

    • J. G. M. Thewissen
    •  & Lisa Noelle Cooper
  2. †School of Biomedical Sciences, Kent State University, Kent, Ohio 44242, USA

    • Lisa Noelle Cooper
  3. ‡Department of Geology and Geophysics, University of Wyoming, Laramie, Wyoming 82071, USA

    • Mark T. Clementz
  4. §Department of Earth Sciences, Indian Institute of Technology, Roorkee 247 667, Uttarakhand, India

    • Sunil Bajpai
  5. Wadia Institute of Himalayan Geology, G.M.S. Road, Dehra Dun 248 001, Uttarakhand 248 001, India

    • B. N. Tiwari


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