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Language-tree divergence times support the Anatolian theory of Indo-European origin


Languages, like genes, provide vital clues about human history1,2. The origin of the Indo-European language family is “the most intensively studied, yet still most recalcitrant, problem of historical linguistics”3. Numerous genetic studies of Indo-European origins have also produced inconclusive results4,5,6. Here we analyse linguistic data using computational methods derived from evolutionary biology. We test two theories of Indo-European origin: the ‘Kurgan expansion’ and the ‘Anatolian farming’ hypotheses. The Kurgan theory centres on possible archaeological evidence for an expansion into Europe and the Near East by Kurgan horsemen beginning in the sixth millennium BP7,8. In contrast, the Anatolian theory claims that Indo-European languages expanded with the spread of agriculture from Anatolia around 8,000–9,500 years bp9. In striking agreement with the Anatolian hypothesis, our analysis of a matrix of 87 languages with 2,449 lexical items produced an estimated age range for the initial Indo-European divergence of between 7,800 and 9,800 years bp. These results were robust to changes in coding procedures, calibration points, rooting of the trees and priors in the bayesian analysis.

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Figure 1: Consensus tree and divergence-time estimates.

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  1. Pagel, M. in Time Depth in Historical Linguistics (eds Renfrew, C., McMahon, A. & Trask, L.) 189–207 (The McDonald Institute for Archaeological Research, Cambridge, UK, 2000)

    Google Scholar 

  2. Gray, R. D. & Jordan, F. M. Language trees support the express-train sequence of Austronesian expansion. Nature 405, 1052–1055 (2000)

    Article  ADS  CAS  Google Scholar 

  3. Diamond, J. & Bellwood, P. Farmers and their languages: the first expansions. Science 300, 597–603 (2003)

    Article  ADS  CAS  Google Scholar 

  4. Richards, M. et al. Tracing European founder lineage in the Near Eastern mtDNA pool. Am. J. Hum. Genet. 67, 1251–1276 (2000)

    Article  CAS  Google Scholar 

  5. Semoni, O. et al. The genetic legacy of Paleolithic Homo sapiens in extant Europeans: a Y chromosome perspective. Science 290, 1155–1159 (2000)

    Article  ADS  Google Scholar 

  6. Chikhi, L., Nichols, R. A., Barbujani, G. & Beaumont, M. A. Y genetic data support the Neolithic Demic Diffusion Model. Proc. Natl Acad. Sci. USA 99, 11008–11013 (2002)

    Article  ADS  CAS  Google Scholar 

  7. Gimbutas, M. The beginning of the Bronze Age in Europe and the Indo-Europeans 3500–2500 B.C. J. Indo-Eur. Stud. 1, 163–214 (1973)

    Google Scholar 

  8. Mallory, J. P. Search of the Indo-Europeans: Languages, Archaeology and Myth (Thames & Hudson, London, 1989)

    Google Scholar 

  9. Renfrew, C. in Time Depth in Historical Linguistics (eds Renfrew, C., McMahon, A. & Trask, L.) 413–439 (The McDonald Institute for Archaeological Research, Cambridge, UK, 2000)

    Google Scholar 

  10. Swadesh, M. Lexico-statistic dating of prehistoric ethnic contacts. Proc. Am. Phil. Soc. 96, 453–463 (1952)

    Google Scholar 

  11. Bergsland, K. & Vogt, H. On the validity of glottochronology. Curr. Anthropol. 3, 115–153 (1962)

    Article  Google Scholar 

  12. Blust, R. in Time Depth in Historical Linguistics (eds Renfrew, C., McMahon, A. & Trask, L.) 311–332 (The McDonald Institute for Archaeological Research, Cambridge, UK, 2000)

    Google Scholar 

  13. Steel, M. A., Hendy, M. D. & Penny, D. Loss of information in genetic distances. Nature 333, 494–495 (1988)

    Article  ADS  Google Scholar 

  14. Swofford, D. L., Olsen, G. J., Waddell, P. J. & Hillis, D. M. in Molecular Systematics (eds Hillis, D., Moritz, C. & Mable, B. K.) 407–514 (Sinauer Associates, Inc, Sunderland, Massachusetts, 1996)

    Google Scholar 

  15. Dixon, R. M. W. The Rise and Fall of Language (Cambridge Univ. Press, Cambridge, UK, 1997)

    Book  Google Scholar 

  16. Metropolis, N., Rosenbluth, A. W., Rosenbluth, M. N., Teller, A. H. & Teller, E. Equations of state calculations by fast computing machines. J. Chem. Phys. 21, 1087–1091 (1953)

    Article  ADS  CAS  Google Scholar 

  17. Huelsenbeck, J. P., Ronquist, F., Nielsen, R. & Bollback, J. P. Bayesian inference of phylogeny and its impact on evolutionary biology. Science 294, 2310–2314 (2001)

    Article  ADS  CAS  Google Scholar 

  18. Huson, D. H. SplitsTree: analyzing and visualizing evolutionary data. Bioinformatics 14, 68–73 (1998)

    Article  CAS  Google Scholar 

  19. Sanderson, M. R8s, Analysis of Rates of Evolution,Version 1.50 (Univ. California, Davis, 2002)

    Google Scholar 

  20. Dyen, I., Kruskal, J. B. & Black, P. FILE IE-DATA1. Available at 〈〉 (1997).

  21. Sanderson, M. J. & Donoghue, M. J. Patterns of variation in levels of homoplasy. Evolution 43, 1781–1795 (1989)

    Article  Google Scholar 

  22. Gamkrelidze, T. V. & Ivanov, V. V. Trends in Linguistics 80: Indo-European and the Indo-Europeans (Mouton de Gruyter, Berlin, 1995)

    Book  Google Scholar 

  23. Rexova, K., Frynta, D. & Zrzavy, J. Cladistic analysis of languages: Indo-European classification based on lexicostatistical data. Cladistics 19, 120–127 (2003)

    Article  Google Scholar 

  24. Ringe, D., Warnow, T. & Taylor, A. IndoEuropean and computational cladistics. Trans. Philol. Soc. 100, 59–129 (2002)

    Article  Google Scholar 

  25. Gkiasta, M., Russell, T., Shennan, S. & Steele, J. Neolithic transition in Europe: the radiocarbon record revisited. Antiquity 77, 45–62 (2003)

    Article  Google Scholar 

  26. Cavalli-Sforza, L. L., Menozzi, P. & Piazza, A. The History and Geography of Human Genes (Princeton Univ. Press, Princeton, 1994)

    MATH  Google Scholar 

  27. Holden, C. J. Bantu language trees reflect the spread of farming across sub-Saharan Africa: a maximum-parsimony analysis. Proc. R. Soc. Lond. B 269, 793–799 (2002)

    Article  Google Scholar 

  28. Barbrook, A. C., Howe, C. J., Blake, N. & Robinson, P. The phylogeny of The Canterbury Tales. Nature 394, 839 (1998)

    Article  ADS  CAS  Google Scholar 

  29. McMahon, A. & McMahon, R. Finding families: Quantitative methods in language classification. Trans. Philol. Soc. 101, 7–55 (2003)

    Article  Google Scholar 

  30. Huelsenbeck, J. P. & Ronquist, F. MRBAYES: Bayesian inference of phylogeny. Bioinformatics 17, 754–755 (2001)

    Article  CAS  Google Scholar 

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We thank S. Allan, L. Campbell, L. Chikhi, M. Corballis, N. Gavey, S. Greenhill, J. Hamm, J. Huelsenbeck, G. Nichols, A. Rodrigo, F. Ronquist, M. Sanderson and S. Shennan for useful advice and/or comments on the manuscript.

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Correspondence to Russell D. Gray.

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Gray, R., Atkinson, Q. Language-tree divergence times support the Anatolian theory of Indo-European origin. Nature 426, 435–439 (2003).

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