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The middle Pliocene gets crowded

New hominin fossils discovered in Ethiopia, dated to between 3.5 million and 3.3 million years ago, suggest that species diversity may have been as high during early human evolution as in later periods. See Article p.483

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Figure 1: Temporal distribution of known hominin species.


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  1. Kimbel, W. H. & Delezene, L. K. Yearb. Phys. Anthropol. 52, 2–48 (2009).

    Article  Google Scholar 

  2. Haile-Selassie, Y. et al. Nature 521, 483–488 (2015).

    Article  CAS  ADS  Google Scholar 

  3. Brunet, M. et al. C. R. Acad. Sci. 322, 907–913 (1996).

    Google Scholar 

  4. Leakey, M. G. et al. Nature 410, 433–440 (2001).

    Article  CAS  ADS  Google Scholar 

  5. Spoor, F., Leakey, M. G. & Leakey, L. N. Phil. Trans. R. Soc. B 365, 3377–3388 (2010).

    Article  Google Scholar 

  6. Guy, F. et al. J. Hum. Evol. 55, 37–47 (2008).

    Article  Google Scholar 

  7. Haile-Selassie, Y. et al. Nature 483, 565–569 (2012).

    Article  CAS  ADS  Google Scholar 

  8. Granger, D. E. et al. Nature (2015).

  9. McPherron, S. P. et al. Nature 466, 857–860 (2010).

    Article  CAS  ADS  Google Scholar 

  10. Harmand, S. et al. Nature 521, 310–315 (2015).

    Article  CAS  ADS  Google Scholar 

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Correspondence to Fred Spoor.

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Spoor, F. The middle Pliocene gets crowded. Nature 521, 432–433 (2015).

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