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First Devonian tetrapod from Asia


The earliest tetrapods (vertebrates with limbs rather than paired fins) date from the Late Devonian Period (370–354 million years ago)1,2 — nine genera have been described, all of which are from the Euramerican supercontinent that comprises Europe, north America and Greenland, apart from a single Gondwanan genus, Metaxygnathus, from Australia3,4,5. Here we report the discovery of the first Devonian tetrapod from Asia, a finding that substantially extends the geographical range of these animals and raises new questions about their dispersal. These forms seem to have achieved worldwide distribution and great taxonomic diversity within a relatively short time.

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Figure 1: A new Devonian tetrapod from China.


  1. Ahlberg, P. E. & Milner, A. R. Nature 368, 507–514 (1994).

    Article  ADS  Google Scholar 

  2. Coates, M. I. & Clack, J. A. Nature 352, 234–236 (1991).

    Article  ADS  Google Scholar 

  3. Campbell, K. S. W. & Bell, M. Alcheringa 1, 369–381 (1977).

    Article  Google Scholar 

  4. Ahlberg, P. E. & Clack, J. A. Trans. R. Soc. Edinb. 89, 11–46 (1998).

    Article  Google Scholar 

  5. Daeschler, E. B. J. Paleontol. 74, 301–308 (2000).

    Article  Google Scholar 

  6. Pan, J. et al. Continental Devonian System of Ningxia and its Biotas (Geological, Beijing, 1987).

    Google Scholar 

  7. Metcalfe, I. in Biogeography and Geological Evolution of SE Asia (eds Hall, R. & Holloway, J. D.) 25–41 (Backbuys, Leiden, The Netherlands, 1998).

    Google Scholar 

  8. Ahlberg, P. E. Nature 373, 420–425 (1995).

    Article  ADS  CAS  Google Scholar 

  9. Ahlberg, P. E., Lukševičs, E. & Mark-Kurik, E. Palaeontology 43, 533–548 (2000).

    Article  Google Scholar 

  10. Milner, A. R. in Palaeozoic Vertebrate Biostratigraphy and Biogeography (ed. Long, J. A.) 324–353 (Belhaven, London, 1993).

    Google Scholar 

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Correspondence to Min Zhu.

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Zhu, M., Ahlberg, P., Zhao, W. et al. First Devonian tetrapod from Asia. Nature 420, 760–761 (2002).

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