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Remains of Homo erectus from Bouri, Middle Awash, Ethiopia


The genesis, evolution and fate of Homo erectus have been explored palaeontologically since the taxon's recognition in the late nineteenth century. Current debate1 is focused on whether early representatives from Kenya and Georgia should be classified as a separate ancestral species (‘H. ergaster’)2,3,4, and whether H. erectus was an exclusively Asian species lineage that went extinct5,6. Lack of resolution of these issues has obscured the place of H. erectus in human evolution. A hominid calvaria and postcranial remains recently recovered from the Dakanihylo Member of the Bouri Formation, Middle Awash, Ethiopia, bear directly on these issues. These 1.0-million-year (Myr)-old Pleistocene sediments contain abundant early Acheulean stone tools and a diverse vertebrate fauna that indicates a predominantly savannah environment. Here we report that the ‘Daka’ calvaria's metric and morphological attributes centre it firmly within H. erectus. Daka's resemblance to Asian counterparts indicates that the early African and Eurasian fossil hominids represent demes of a widespread palaeospecies. Daka's anatomical intermediacy between earlier and later African fossils provides evidence of evolutionary change. Its temporal and geographic position indicates that African H. erectus was the ancestor of Homo sapiens.

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Figure 1: Maps and generalized sections.
Figure 2: Views of the Daka calvaria and cladograms representing majority rule (a) and strict consensus (b) of the 75 most parsimonious phylogenies generated by PAUP 4.0b8 (Sinauer).


  1. Delson, E. Palaeobiology and age of African Homo erectus. Nature 316, 762–763 (1985).

    ADS  CAS  Article  Google Scholar 

  2. Walker, A. C. & Leakey, R. E. F. The Nariokotome Homo erectus Skeleton (Harvard Univ. Press, Cambridge, Massachusetts, 1993).

    Book  Google Scholar 

  3. Wood, B. A. Koobi Fora Research Project Vol. 4 (Clarendon Press, Oxford, 1991).

    Google Scholar 

  4. Gabunia, L., Vekua, A. & Lordkipanidze, D. New human fossils from Dmanisi, eastern Georgia. Archaeol. Ethnol. Anthropol. Eurasia 2(6), 128–139 (2001).

    Google Scholar 

  5. Andrews, P. J. An alternative interpretation of the characters used to define Homo erectus. Courier Forschunginstitut Senckenberg 69, 167–175 (1984).

    Google Scholar 

  6. Groves, C. P. A Theory of Human and Primate Evolution (Clarendon Press, Oxford, 1989).

    Google Scholar 

  7. DeHeinzelin, J. et al. Environment and behavior of 2.5-million-year-old Bouri hominids. Science 284, 625–629 (1999).

    ADS  CAS  Article  Google Scholar 

  8. DeHeinzelin, J., Clark, J. D., Schick, K. D. & Gilbert, W. H. (eds) The Acheulean and the Plio-Pleistocene Deposits of the Middle Awash Valley, Ethiopia (Royal Museum of Central Africa, Tervuren, Belgium; Ann. Sci. Geol., 104 (2000)).

    Google Scholar 

  9. Kalb, J. et al. Geology and stratigraphy of Neogene deposits, Middle Awash Valley, Ethiopia. Nature 298, 17–25 (1982).

    ADS  Article  Google Scholar 

  10. Vrba, E. S. New fossils of Alcelaphini and Caprinae (Bovidae: Mammalia) from Awash, Ethiopia, and phylogenetic analysis of Alcelaphini. Palaeont. Afr. 34, 127–198 (1997).

    Google Scholar 

  11. Abbate, E. et al. A one-million-year-old Homo cranium from the Danakil (Afar) depression of Eritrea. Nature 393, 458–460 (1998).

    ADS  CAS  Article  Google Scholar 

  12. Brauer, G. & Mbua, E. Homo erectus features used in cladistics and their variability in Asian and African hominids. J. Hum. Evol. 22, 79–108 (1992).

    Article  Google Scholar 

  13. Kennedy, G. E. On the autapomorphic traits of Homo erectus. J. Hum. Evol. 20, 375–412 (1991).

    Article  Google Scholar 

  14. Kramer, A. Human taxonomic diversity in the Pleistocene: Does Homo erectus represent multiple hominid species? Am. J. Phys. Anthropol. 91, 161–171 (1993).

    CAS  Article  Google Scholar 

  15. Harrison, T. in Species, Species Concepts, and Primate Evolution (eds Kimbel, W. H. & Martin, L. B.) 345–371 (Plenum, New York, 1993).

    Book  Google Scholar 

  16. Turner, A. & Chamberlain, A. Speciation, morphological change and the status of African Homo erectus. J. Hum. Evol. 18, 115–130 (1989).

    Article  Google Scholar 

  17. Lovejoy, C. O., Cohn, M. J. & White, T. D. Morphological analysis of the mammalian postcranium: A developmental perspective. Proc. Natl Acad. Sci. USA 96, 13247–13252 (1999).

    ADS  CAS  Article  Google Scholar 

  18. Trinkaus, E. Cladistics and the hominid fossil record. Am. J. Phys. Anthropol. 83, 1–11 (1990).

    CAS  Article  Google Scholar 

  19. Howell, F. C. Paleo-demes, species clades, and extinctions in the Pleistocene hominin record. J. Anthropol. Res. 55, 191–243 (1999).

    Article  Google Scholar 

  20. Martinez, I. & Arsuaga, J. L. The temporal bones from Sima de los Huesos Middle Pleistocene site (Sierra de Atapuerca, Spain). A phylogenetic approach. J. Hum. Evol. 33, 283–318 (1997).

    CAS  Article  Google Scholar 

  21. Rightmire, G. P. The Evolution of Homo erectus: Comparative Anatomical Studies of an Extinct Human Species (Cambridge Univ. Press, Cambridge, 1990).

    Book  Google Scholar 

  22. Rightmire, G. P. Evidence from facial morphology for similarity of Asian and African representatives of Homo erectus. Am. J. Phys. Anthropol. 106, 61–85 (1998).

    CAS  Article  Google Scholar 

  23. Aguirre, E. Poor fossil record and major changes around 1 MaBP. Hum. Evol. 15, 51–62 (2000).

    Article  Google Scholar 

  24. Larick, R. et al. Early Pleistocene 40Ar/39Ar ages for Bapang Formation hominins, Central Jawa, Indonesia. Proc. Natl Acad. Sci. USA 98, 4866–4871 (2001).

    ADS  CAS  Article  Google Scholar 

  25. Olsen, J. W. New light on the earliest occupation of east Asia. Anthropologie 37, 89–96 (1999).

    Google Scholar 

  26. Rightmire, G. P. The human cranium from Bodo, Ethiopia: Evidence for speciation in the Middle Pleistocene. J. Hum. Evol. 31, 21–39 (1996).

    Article  Google Scholar 

  27. Schwartz, J. H. Taxonomy of the Dmanisi crania. Science 289, 55 (2000).

    CAS  Article  Google Scholar 

  28. Ambrose, S. H. Paleolithic technology and human evolution. Science 291, 1748–1753 (2001).

    ADS  CAS  Article  Google Scholar 

  29. Bar-Yosef, O. Lower Paleolithic sites in South-western Asia: Evidence for ‘Out of Africa’ movements. Anthropologie 37, 51–69 (1999).

    Google Scholar 

  30. Clarke, R. Out of Africa and back again. Int. J. Anthropol. 15, 185–189 (2001).

    Article  Google Scholar 

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This paper is dedicated to the late J. Desmond Clark who initiated this research at Bouri in 1981. We thank A. Almquist, A. Asfaw, M. Asnake, T. Assebework, D. Brill, D. DeGusta, J. DeHeinzelin, A. Getty, Y. Haile-Selassie, B. Latimer, Ç. Pehlevan, K. Schick, S. Simpson, P. Snow, G. Suwa and Y. Zeleke for fieldwork and analytical studies. Electron microprobe and other support from the Earth Environmental Sciences Division, Los Alamos National Laboratory. Thanks to G. Suwa, D. DeGusta, C. Feibel, F. C. Howell, C. O. Lovejoy, F. Bibi, T. Stidham, J. Parham, J.-R. Boisserie and H. Saegusa for thoughtful review and/or assistance. We thank the Ministry of Youth, Sports and Culture, the Authority for Research and Conservation of the Cultural Heritage, and the National Museum of Ethiopia for permissions, and the Afar Regional Government and the Afar people of the Middle Awash, particularly H. Elema. Many additional individuals contributed. This research was supported by the NSF (USA) and the Institute of Geophysics and Planetary Physics (University of California at Los Alamos National Laboratory), with additional contributions by the Graduate School and Hampton Fund for International Initiatives, Miami University.

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Correspondence to Berhane Asfaw.

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Asfaw, B., Gilbert, W., Beyene, Y. et al. Remains of Homo erectus from Bouri, Middle Awash, Ethiopia. Nature 416, 317–320 (2002).

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