SEEDS of some pines, such as Jeffrey pine (Pinus jeffreyi), can be stored in air-tight jars at room temperature for a long time. Even after ten years of storage one may expect as much as 40 per cent of viable seed. When the seeds of Jeffrey pine are sown in a greenhouse they germinate rapidly and abundantly. Many other pines, as, for example, sugar pine (Pinus lambertiana), have short-lived seed. Under ordinary storage conditions sugar pine seeds lose their viability rapidly, and after five years of storage their germination is usually nil. At 5° C. the viability of sugar pine seed is maintained for a long time; in one case, after eight years of cold storage, germination amounted to 86 per cent of the original. Normally seeds of sugar pine sown in a greenhouse either fail to germinate completely or give a very small percentage of germination. When, however, the seeds are chilled in some moist medium for three months at 5° C, they germinate as well as those of Jeffrey pine. This prolonged period at a low temperature, necessary for germination of refractory seeds, can be designated as the period of incipient germination.
Meyerhof, Otto, "Chemical Dynamics of Life Phaenomena" (Philadelphia and London, J. B. Lippincott Co., 1924).
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MIROV, N. Possible Relation of Linolenic Acid to the Longevity and Germination of Pine Seed. Nature 154, 218–219 (1944). https://doi.org/10.1038/154218a0
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