Takayuki Nishizaka1, 2, 7, Kazuhiro Oiwa1, Hiroyuki Noji3, Shigeki Kimura1, Eiro Muneyuki4, Masasuke Yoshida4, 5
& Kazuhiko Kinosita Jr61
Kansai Advanced Research Center, Protein Biophysics Group, Iwaoka 588-2, Nishi-ku, Kobe 651-2492, Japan.
2
Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency, Honcho 4-1-8, Kawaguchi, Saitama 332-0012, Japan.
3
Institute of Industrial Science, University of Tokyo, Tokyo 153-8505, Japan.
4
Chemical Resources Laboratory, Tokyo Institute of Technology, Yokohama 226-8503, Japan.
5
ATP system, Exploratory Research for Advanced Technology (ERATO), Japan Science and Technology Agency, Yokohama 226-0026, Japan.
6
Center for Integrative Bioscience, Okazaki National Research Institutes, Myodaiji, Okazaki 444-8585, Japan.
7
Present address: Department of Physics, Gakushuin University, Mejiro 1-5-1, Toshima-ku, Tokyo 171-8588, Japan.
Correspondence should be addressed to Takayuki Nishizaka takayuki.nishizaka@gakushuin.ac.jp
subunit is powered by ATP hydrolysis in three catalytic sites arranged 120° apart around 
80° rotation of 
