Nature Reviews Molecular Cell Biology 7, 829-832 (November 2006) | doi:10.1038/nrm2023

EssayBack to the future: education for systems-level biologists

Ned Wingreen1 & David Botstein1  About the authors


We describe a graduate course in quantitative biology that is based on original path-breaking papers in diverse areas of biology; each of these papers depends on quantitative reasoning and theory as well as experiment. Close reading and discussion of these papers allows students with backgrounds in physics, computational sciences or biology to learn essential ideas and to communicate in the languages of disciplines other than their own.

Author affiliations

  1. Ned Wingreen is at the Department of Molecular Biology, and David Botstein is at the Lewis-Sigler Institute and the Department of Molecular Biology, Princeton University, Princeton, New Jersey 08544, USA.

Correspondence to: Email:

Published online 20 September 2006


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