1
James J. Bull is in the Section of Integrative Biology and Institute for Cellular and Molecular Biology, University of Texas, Austin, Texas 78712-0253, USA. bull@bull.biosci.utexas.edu
2
Sarah P. Otto is in the Department of Zoology, University of British Columbia, Vancouver, British Columbia V6T 1Z4, Canada.
The first empirical test of an evolutionary theory provides support for a mutational landscape model underlying the process of adaptation. The study shows that it is possible to predict at least the first step in an adaptive walk and also shows the importance of incorporating mutation bias in the fitness effects of mutations.
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