Hand-operated pump would have kept catch alive during long trips.
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References
Beltrame, C., Gaddi, D. & Parizzi, S. Int. J. Naut. Archaeol. doi:10.1111/j.1095-9270.2011.00317.x (2011)
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Marchant, J. Roman ship had on-board fish tank. Nature (2011). https://doi.org/10.1038/news.2011.335
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DOI: https://doi.org/10.1038/news.2011.335