Miniature transponders show pattern among the erratic fluttering.
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References
CantE. T., SmithA. D., ReynoldsD. R. & OsborneJ. L. et al. Proc. Roy. Soc. B., doi:10.1098/rspb.2004.3002 (2004).
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Rothamsted Research
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Giles, J. Radar reveals purpose in butterfly flights. Nature (2005). https://doi.org/10.1038/news050404-4
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DOI: https://doi.org/10.1038/news050404-4