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The tomato (Solanum lycopersicum) is a major crop plant worldwide and a model organism for the Solanaceae family of flowering plants. It is widely used for the study of fruit biology and disease resistance. A high-quality genome sequence of a domesticated tomato — the Heinz 1706 cultivar — has now been determined, together with a draft sequence for its closest wild relative, the 'currant tomato' Solanum pimpinellifolium. Comparative genomics identifies only 0.6% difference between the two, but a divergence of more than 8% from the potato, Solanum tuberosum, which was sequenced last year. The tomato and S. pimpinellifolium sequences also record the bottlenecks that have narrowed genetic diversity: domestication in the Americas; the export of a few genotypes to Europe in the sixteenth century; and centuries of intensive breeding. Cover: natu/Shutterstock.