Department of Zoology, University of Melbourne, Melbourne, Victoria 3010, Australia
Andrew F. Hugall &
Present address: Sciences Department, Melbourne Museum, Melbourne, Victoria 3053, Australia.
Andrew F. Hugall
A.F.H. constructed phylogenies, conducted diversification analyses, wrote the Methods, Supplementary Information and edited the main manuscript. D.S.-F. conceived and funded the project, wrote the main manuscript and edited remaining sections. Both authors contributed to interpretation of results.
Competing financial interests
The authors declare no competing financial interests.