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Nature 438, E4 (17 November 2005) | doi:10.1038/nature04356; Published online 16 November 2005

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Origin of flight: Could 'four-winged' dinosaurs fly? (Reply)

Zhonghe Zhou1 & Fucheng Zhang1

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Padian and Dial1 challenge our view that the evolution of flight involved a four-winged stage. This disagreement stems from our different views on the origin of bird flight and from the methodology we use to analyse functional morphology in the non-avian theropod Microraptor2 and in an enantiornithine bird3 from the Early Cretaceous period in China.

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