 

Mean Fitness Increases when Fitnesses are Additive W. J. EWENS Department of Mathematics, La Trobe University, Melbourne. FISHER'S fundamental theorem of natural selection^{1} states that for a random mating population with fitnesses dependent on the genetic constitution at a single locus, the mean fitness of the population increases with time. This is true for an arbitrary number of possible alleles at the locus and for arbitrary fitness values.
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