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Nature 422, 606-608 (10 April 2003) | doi:10.1038/nature01526;

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N2 production by the anammox reaction in the anoxic water column of Golfo Dulce, Costa Rica

In oxygen-depleted zones of the open ocean, and in anoxic basins and fjords, denitrification (the bacterial reduction of nitrate to give N2) is recognized as the only significant process converting fixed nitrogen to gaseous N2. Primary production in the oceans is often limited by the availability of fixed nitrogen such as ammonium or nitrate, and nitrogen-removal processes consequently affect both ecosystem function and global biogeochemical cycles.

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