A motor protein can be made to walk in either direction along a filamentous track by adjusting the concentration of calcium ions in the surrounding solution.
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Walter, W., Diez, S. Myosin shifts into reverse gear. Nature Nanotech 7, 213–214 (2012). https://doi.org/10.1038/nnano.2012.49
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DOI: https://doi.org/10.1038/nnano.2012.49